Calcium (Ca2+) signaling is integral to nearly all aspects of plant biology, including development and responses to biotic and abiotic stresses. It operates through two main layers: the generation of Ca2+ signals and their decoding by Ca2+-binding proteins, which act early in diverse signaling pathways. The system exhibits remarkable robustness and versatility, largely due to its network-like organization. While fundamental principles of Ca2+ signaling were initially established in noncrop model organisms